2012 in archosaur paleontology

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The year 2012 in Archosaur paleontology was eventful. Archosaurs include the only living dinosaur group — birds — and the reptile crocodilians, plus all extinct dinosaurs, extinct crocodilian relatives, and pterosaurs. Archosaur palaeontology is the scientific study of those animals, especially as they existed before the Holocene Epoch began about 11,700 years ago. The year 2012 in paleontology included various significant developments regarding archosaurs.

This article records new taxa of fossil archosaurs of every kind that have been described during the year 2012, as well as other significant discoveries and events related to paleontology of archosaurs that occurred in the year 2012.

Pseudosuchians

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Aegisuchus[5]

Gen. et sp. nov

Valid

Holliday & Gardner

Cenomanian

Kem Kem Formation

 Morocco

An aegyptosuchid eusuchian.

Aegisuchus witmeri.

Aetobarbakinoides[6]

Gen. et sp. nov

Valid

Desojo, Ezcurra & Kischlat

Late Carnian or early Norian

Santa Maria Formation

 Brazil

An aetosaur.

Apachesuchus[7]

Gen. et sp. nov

Valid

Spielmann & Lucas

Late Triassic

 United States

An aetosaur. The type species is Apachesuchus heckerti.

Barreirosuchus[8]

Gen. et sp. nov

Valid

Iori & Garcia

Turonian to Santonian

Adamantina Formation

 Brazil

A trematochampsid. The type species is Barreirosuchus franciscoi.

Borealosuchus threeensis[9]

Sp. nov

Valid

Brochu, Parris, Grandstaff, Denton & Gallagher

Maastrichtian or Danian

 USA

A crocodilian from New Jersey.

Bystrowisuchus[10]

Gen. et sp. nov

Valid

Sennikov

Early Triassic

 Russia

A ctenosauriscid. The type species is Bystrowisuchus flerovi.

Crocodylus thorbjarnarsoni[11]

sp nov

Valid

Brochu & Storrs

Pliocene to Pleistocene

 Kenya

A species of Crocodylus.

Crocodylus thorbjarnarsoni.

Gasparinisuchus[12]

Gen. et sp. nov

Valid

Martinelli et al.

Late Cretaceous

 Argentina

A peirosaurid. The type species is Gasparinisuchus peirosauroides.

Newly named basal dinosauriforms

Name Novelty Status Authors Age Unit Location Notes Images

Diodorus[13]

Gen. et sp. nov

Valid

Kammerer, Nesbitt, & Shubin

?Carnian-Norian

Timezgadiouine Formation

 Morocco

A silesaurid.

Non-avian dinosaurs

Research

New taxa

Name Novelty Status Authors Age Unit Location Notes Images

Alnashetri[54]

Gen. et sp. nov

Valid

Makovicky, Apesteguía & Gianechini

Late Cretaceous (Cenomanian or Turonian)

Candeleros Formation

 Argentina

A coelurosaurian, a probable relative of alvarezsaurids. The type species is Alnashetri cerropoliciensis.

Astrophocaudia[55]

Gen. et sp. nov

Valid[56]

D’Emic

Early Cretaceous

Trinity Group

 United States

A titanosauriform sauropod, a member of the clade Somphospondyli. The type species is Astrophocaudia slaughteri.

Batyrosaurus[57]

Gen. et sp. nov

Valid

Godefroit et al.

Late Cretaceous

 Kazakhstan

A basal hadrosauroid. The type species is Batyrosaurus rozhdestvenskyi.

Bicentenaria[58]

Gen. et sp. nov

Valid

Novas et al.

Cenomanian

Candeleros Formation

 Argentina

A coelurosaur. The type species is Bicentenaria argentina.

Bonapartenykus[59]

Gen. et sp. nov

Valid

Agnolin, Powell, Novas, & Kundrát

Campanian/Maastrichtian

Allen Formation

 Argentina

An alvarezsaurid.

Comahuesaurus[60]

Gen. et sp. nov

Valid

Carballido et al.

Early Cretaceous (Aptian or Albian)

Lohan Cura Formation

 Argentina

A rebbachisaurid sauropod. The type species is Comahuesaurus windhauseni.

Coronosaurus[61]

Gen. et comb. nov

Valid

Ryan, Evans & Shepherd

Late Cretaceous

Oldman Formation

 Canada

A centrosaurine ceratopsian, a new genus for "Centrosaurus" brinkmani Ryan & Russell (2005).

Darwinsaurus[62]

Gen. et sp. nov

Valid

Paul

Early Cretaceous (early Valanginian)

Wadhurst Clay Formation

 United Kingdom

A styracosternan ornithopod. The type species is Darwinsaurus evolutionis, and the holotype is an associated skeleton that includes material catalogued under numbers NHMUK R1831 (a dentary), R1833, R1835 and R1836;[62] this skeleton is assigned by Norman (2010) to Hypselospinus fittoni.[63]

Elaltitan[64]

Gen. et sp. nov

Valid

Mannion & Otero

Middle Cenomanian to Turonian

Bajo Barreal Formation

 Argentina

A lithostrotian titanosaur. The type species is Elaltitan lilloi.

Eoabelisaurus[65]

Gen. et sp. nov

Valid

Pol & Rauhut

Aalenian or Bajocian

Cañadon Asfalto Formation

 Argentina

An abelisaurid. The type species is Eoabelisaurus mefi.

Gideonmantellia[66]

Gen. et sp. nov

Valid

Ruiz-Omeñaca et al.

Barremian

Camarillas Formation

 Spain

A basal ornithopod. The type species is Gideonmantellia amosanjuanae.

Gryphoceratops[67]

Gen. et sp. nov

Valid

Ryan, Evans, Currie, Brown, & Brinkman

Late Santonian

Milk River Formation

 Canada

A leptoceratopsid ceratopsian.

Hexing[68]

Gen. et sp. nov

Valid

Jin, Chen & Godefroit

Early Valanginian to early Barremian

Yixian Formation

 China

A basal ornithomimosaur. The type species is Hexing qingyi.

Huehuecanauhtlus[69]

Gen. et sp.

Valid

Ramírez-Velasco et al.

Santonian

 Mexico

A basal hadrosauroid. The type species is Huehuecanauhtlus tiquichensis.

Huehuecanauhtlus.

Huxleysaurus[62]

Gen. et com. nov

Valid

Paul

Early Cretaceous (early Valanginian)

Wadhurst Clay Formation

 United Kingdom

A styracosternan ornithopod, a new genus for "Iguanodon" hollingtoniensis Lydekker (1889);[62] this species is considered by Norman (2010) to be a junior synonym of Hypselospinus fittoni.[63]

Ichthyovenator[70]

Gen. et sp. nov

Valid

Allain et al.

Late Early Cretaceous

Savannakhet Basin

 Laos

A spinosaurid. The type species is Ichthyovenator laosensis.

Kaatedocus[71]

Gen. et sp. nov

Valid[72]

Tschopp & Mateus

Late Jurassic (Kimmeridgian)

Morrison Formation

 United States

A diplodocine diplodocid sauropod. The type species is Kaatedocus siberi.

Kundurosaurus[73]

Gen. et sp. nov

Valid

Godefroit, Bolotsky & Lauters

Late Cretaceous, possibly late Maastrichtian

Udurchukan Formation

 Russia

A saurolophine hadrosaurid. The type species is Kundurosaurus nagornyi.

Kundurosaurus.

Lapampasaurus[74]

Gen. et sp. nov

Valid

Coria, Riga & Casadío

Late Cretaceous (late Campanian or early Maastrichthian)

Allen Formation

 Argentina

A hadrosaurid. The type species is Lapampasaurus cholinoi.

Latirhinus[75]

Gen. et sp. nov

Valid

Prieto-Márquez & Brañas

Late Cretaceous (late Campanian)

Cerro del Pueblo Formation

 Mexico

A saurolophine hadrosaurid. The type species is Latirhinus uitstlani.

Magnapaulia[76]

Gen. et comb. nov

Valid

Prieto-Márquez, Chiappe & Joshi

Late Campanian

El Gallo Formation

 Mexico

A lambeosaurine hadrosaurid, a new genus for "Lambeosaurus" laticaudus (Morris, 1981).

Magnapaulia.

Mantellodon[62]

Gen. et sp. nov

Valid

Paul

Early Cretaceous (early Aptian)

Lower Greensand Formation

 United Kingdom

A styracosternan ornithopod. The type species is Mantellodon carpenteri and the holotype is the specimen NHMUK R3741 consisting of partial associated skeleton (also known as Gideon Mantell's "Mantel-piece")[62]

Martharaptor[77]

Gen. et sp. nov

Valid

Senter, Kirkland & DeBlieux

Early Aptian

Cedar Mountain Formation

 United States

A possible therizinosauroid. The type species is Martharaptor greenriverensis.

Martharaptor.

Microraptor hanqingi[78]

sp nov

Valid

Gong et al.

Early Cretaceous

 China

A species of Microraptor. Its status as a distinct species is disputed, with Pei et al. (2014) considering the holotype specimen's morphology not to be distinguishable from other Microraptor specimens.[79]

Mochlodon vorosi[80]

Sp. nov

Valid

Ősi et al.

Santonian

Csehbánya Formation

 Hungary

A rhabdodontid ornithopod, a species of Mochlodon.

Ningyuansaurus[81]

Gen. et sp. nov

Valid

Ji et al.

Early Cretaceous

Yixian Formation

 China

An oviraptorosaur. The type species is Ningyuansaurus wangi.

Ostafrikasaurus[82]

Gen. et sp. nov

Valid

Buffetaut

Late Jurassic

Tendaguru Formation

 Tanzania

A spinosaurid. The type species is Ostafrikasaurus crassiserratus.

Pachyrhinosaurus perotorum[83]

sp nov

Valid

Fiorillo & Tykoski

Maastrichtian

Prince Creek Formation

 USA

Third known species of Pachyrhinosaurus.

Pegomastax[84]

Gen. et sp. nov

Valid

Sereno

Hettangian to Sinemurian

Elliot Formation

 South Africa

A heterodontosaurid. The type species is Pegomastax africana.

Philovenator[85]

Gen. et sp. nov

Valid

Xu et al.

Campanian

Wulansuhai Formation

 China

A troodontid. Its type species is Philovenator curriei.

Proa[86]

Gen. et sp. nov

Valid

McDonald et al.

Early Cretaceous (early Albian)

Escucha Formation

 Spain

An iguanodont ornithopod closely related to Iguanodon and the clade Hadrosauroidea. The type species is Proa valdearinnoensis.

Rugocaudia[87]

Gen. et sp. nov

Valid

Woodruff

Aptian or Albian

Cloverly Formation

 USA

A titanosauriform sauropod. The type species is Rugocaudia cooneyi.

Sciurumimus[88]

Gen. et sp. nov

Valid

Rauhut et al.

Upper Kimmeridgian

 Germany

A megalosauroid. The type species is Sciurumimus albersdoerferi.

Sciurumimus.

Unescoceratops[67]

Gen. et sp. nov

Valid

Ryan, Evans, Currie, Brown, & Brinkman

Late Campanian

Dinosaur Park Formation

 Canada

A leptoceratopsid ceratopsian.

Xenoceratops[61]

Gen. et sp. nov

Valid

Ryan, Evans & Shepherd

Late Cretaceous (middle Campanian)

Foremost Formation

 Canada

A centrosaurine ceratopsian. The type species is Xenoceratops foremostensis.

Yueosaurus[89]

Gen. et sp. nov

Valid

Zheng, Jin, Shibata, Azuma, & Yu

Aptian-Cenomanian

Liangtoutang Formation

 China

A basal ornithopod.

Yurgovuchia[90]

Gen. et sp. nov

Valid

Senter et al.

Early Cretaceous, possibly Barremian

Cedar Mountain Formation

 USA

A dromaeosaurid. The type species is Yurgovuchia doellingi.

Yurgovuchia.

Yutyrannus[91]

Gen. et sp. nov

Valid

Xu et al.

Early Cretaceous

Yixian Formation

 China

A basal tyrannosauroid. The type species is Yutyrannus huali.

Yutyrannus.

Newly named birds

Name Novelty Status Authors Age Unit Location Notes Images

Aegolius gradyi [92]

Sp. nov.

Valid

Storrs L. Olson

Pleistocene to Holocene

 Bermuda

A Strigidae.

Aegypius jinniushanensis [93]

Sp. nov.

Valid

Zhang Zihui

Huang Yunping

Helen F. James

Hou Lianhai

Middle Pleistocene

 China

An Accipitridae.

Aix praeclara [94]

Sp. nov.

Valid

Nikita V. Zelenkov

Evgeny N. Kurochkin

Middle Miocene

Oshin Formation

 Mongolia

An Anatidae.

Alauda xerarvensis [95]

Sp. nov.

Valid

Zlatozar Boev

Late Pliocene

Middle Villafranchian

 Bulgaria

An Alaudidae.

Anchigyps voorhiesi [96]

Gen. nov. et Sp. nov.

Valid

Zhang Zihui

Alan Feduccia

Helen F. James

Late Miocene.

late Clarendonian

Ash Hollow Formation

 USA:

 Nebraska

An Accipitridae close in morphology to the Old World Vultures. The type species of the new genus.

Anhinga walterbolesi [97]

Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene or Early Miocene

Etadunna Formation,

24-26 Ma

 Australia:

 South Australia

An Anhingidae.

Anthus antecedens [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 7-8

 Hungary

A Motacillidae.

Athene vallgornerensis [100]

Sp. nov.

Valid

Carmen Guerra

Pere Bover

Josep A. Alcover

Early Pleistocene

Cave deposits

 Spain:

 Majorca

A Strigidae.

Australopicus nelsonmandelai [101]

Gen. nov. et Sp. nov.

Valid

Albrecht Manegold

Antoine Louchart

Early Pliocene

Varswater Formation

 South Africa

A Picidae, this is the type species of the new genus.

Avolatavis tenens [102]

Gen. nov. et Sp. nov.

Valid

Daniel T. Ksepka

Julia A. Clarke

Eocene

Green River Formation

 USA:

 Wyoming

A stem parrot, a member of Pan-Psittaciformes. The type species of the new genus.

Becassius charadriioides [103]

Gen. nov. et Sp. nov.

Valid

Vanesa L. De Pietri

Gerald Mayr

Early Miocene

Saint-Gérard-le-Puy,

MN 2a

 France

A Scolopacidae or a relative of the Scolopacidae. The type species of the new genus.

Bombycilla hamori [98] [99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early Miocene

MN 5

 Hungary

A Bombycillidae.

Brodavis americanus [104]

Gen. nov. et Sp. nov.

Valid

Larry D. Martin

Evgeny N. Kurochkin

Tim T. Tokaryk

Late Cretaceous,

Maastrichtian

Frenchman Formation

 Canada:

 Saskatchewan

A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012, this is the type species of the new genus.

Brodavis baileyi [104]

Sp. nov.

Valid

Larry D. Martin

Evgeny N. Kurochkin

Tim T. Tokaryk

Late Cretaceous,

Maastrichtian

Hell Creek Formation

 USA:

 South Dakota

A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012.

Brodavis mongoliensis [104]

Sp. nov.

Valid

Larry D. Martin

Evgeny N. Kurochkin

Tim T. Tokaryk

Late Cretaceous,

Maastrichtian

Nemegt Formation

 Mongolia

A Hesperornithiformes Fürbringer, 1888, Brodavidae Martin, Kurochkin et Tokaryk, 2012.

Certhia janossyi [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Late Miocene

MN 9

 Hungary

A Certhiidae.

Chenoanas deserta [105]

Gen. nov. et Sp. nov.

Valid

Nikita V. Zelenkov

Middle Miocene

Oshin Formation

 Mongolia

An Anatidae, this is the type species of the new genus.

Cinclus major [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early Miocene

MN 5

 Hungary

A Cinclidae.

Circaetus rhodopensis [106]

Sp. nov.

Valid

Zlatozar Boev

Late Miocene

Tulorian = Late Meotian,

Late MN 11, ca. 7,5 Mya

 Bulgaria

An Accipitridae.

Crypturellus reai [107]

Sp. nov.

Valid

Robert M. Chandler

Miocene (Santacrucian)

Santa Cruz Formation

 Argentina

A Tinamidae.

?Elorius limosoides [103]

Sp. nov.

Valid

Vanessa L. De Pietri

Gerald Mayr

Early Miocene

Saint-Gérard-le-Puy,

MN 2a

 France

A Scolopacidae, possibly a species of Elorius.

Emberiza bartkoi [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early Miocene

MN 5

 Hungary

An Emberizidae.

Eremarida xerophila [95]

Gen. nov. et Sp. nov.

Valid

Zlatozar Boev

Late Miocene

Gnilyane Formation

 Bulgaria

An Alaudidae, this is the type species of the new genus.

Eremophila prealpestris [95]

Sp. nov.

Valid

Zlatozar Boev

Late Pliocene

Middle Villafranchian,

MP 17, 2,23-2,40 Mya

 Bulgaria

An Alaudidae.

Erithacus horusitzkyi [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 7/8

 Hungary

A Muscicapidae, an Erithacus robin.

Galerida bulgarica [95]

Sp. nov.

Valid

Zlatozar Boev

Late Pliocene

Middle Villafranchian

 Bulgaria

An Alaudidae.

Galerida cserhatensis [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early Miocene

MN 5

 Hungary

An Alaudidae.

Heteroanser vicinus [108]

Gen. nov. et Comb. nov.

Valid

Nikita V. Zelenkov

Late Miocene

Hyargas Nuur Formation

 Mongolia

An Anatidae a new genus for "Heterochen" vicinus Kurochkin, 1976.

Hirundo selengenica [109]

Sp. nov.

Valid

Nikita V. Zelenkov

Evgeny N. Kurochkin

Late Pliocene

Chikoi Formation

 Mongolia

A Hirundinidae.

Jeholornis palmapenis [110]

Sp. nov.

Valid

Jingmai K. O'Connor

Sun Chengkai

Xu Xing

Wang Xiaolin

Zhou Zhonghe

Early Aptian

Jiufotang Formation

 China

A Jeholornithidae Zhou et Zhang, 2006.

Kairuku grebneffi [111]

Sp. nov.

Valid

Daniel T. Ksepka

R. Ewan Fordyce

Tatsuro Ando

Craig M. Jones

Oligocene

Dutroonian

 New Zealand

A Spheniscidae.

Kairuku waitaki [111]

Gen. nov. et Sp. nov

Valid

Daniel T. Ksepka

R. Ewan Fordyce

Tatsuro Ando

Craig M. Jones

Oligocene

Late Whaingaroan or Early Duntroonian

 New Zealand

A Spheniscidae, the type species of the genus.

Lanius schreteri [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 7-8

 Hungary

A Laniidae.

Leptoptilos lüi [112]

Sp. nov.

Not valid

Zhang Zihui

Huang Yunping

Helen F. James

Hou Lianhai

Middle Pleistocene

Cave deposits

 China

A Ciconiidae, the name is not valid because of the "ü" in it, that must, of course be a plain "u".

Lullula balcanica [95]

Sp. nov.

Valid

Zlatozar Boev

Late Pliocene

Middle Villafranchian

 Bulgaria

An Alaudidae.

Lullula neogradensis [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 7/8

 Hungary

An Alaudidae.

Lullula slivnicensis [95]

Sp. nov.

Valid

Zlatozar Boev

Late Pliocene

Early Late Villafranchian,

MN 18

 Bulgaria

An Alaudidae, a relative of the woodlark.

Luscinia praeluscinia [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early Miocene

MN 5

 Hungary

A Muscicapidae.

Lutetodontopteryx tethyensis [113]

Gen. nov. et Sp. nov.

Valid

Gerald Mayr

Evgenij Zvonok

Middle Eocene

Early Lutetian

 Ukraine

A Pelagornithidae, this is the type species of the new genus.

Melanocorypha donchevi [95]

Sp. nov.

Valid

Zlatozar Boev

Late Pliocene

Middle Villafranchian

 Bulgaria

An Alaudidae.

Melanocorypha serdicensis [95]

Sp. nov.

Valid

Zlatozar Boev

Late Miocene

Gnilyane Formation,

Middle Late Pontian

 Bulgaria

An Alaudidae.

Mioquerquedula minutissima [94]

Gen. nov. et Sp. nov.

Valid

Nikita V. Zelenkov

Evgeny N. Kurochkin

Middle Miocene, possibly also Late Miocene

Gobi-Altai Aimag,

Shargyn Govi,

Sharga location

 Mongolia

An Anatidae, this is the type species of the new genus, the new genus also contains "Anas" velox Milne-Edwards, 1868 as a Comb. nov. from France, Germany, Romania and ?Hungary.

Muscicapa leganyii [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 7/8

 Hungary

A Muscicapidae.

Otus mauli [114]

Sp. nov.

Valid

Carlos Rando

Harald Pieper

Josep A. Alcover

Storrs L. Olson

Quaternary

 Portugal:

 Madeira

A Strigidae.

?Parvelorius calidris [103]

Gen. nov. et Sp. nov.

Valid

Vanessa L. De Pietri

Gerald Mayr

Early Miocene

MN 2a

 France

A Scolopacidae, the new genus is erected for the species "Tringa" gracilis Milne-Edwards, 1868, the type species of the new genus and makes a Com. nov. with it; genus may also contain this new species.

Phylloscopus miocaenicus [98]

Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 6-8

 Hungary

A Phylloscopidae.

Picavus litencicensis [115]

Gen. nov. et Sp. nov.

Valid

Gerald Mayr

Růžena Gregorová

Early Oligocene

Menilite Formation

 Czech Republic

A stem group representative of the Piciformes, placed in a new family Picavidae Mayr et Gregorová, 2012. it is the type species of the new genus.

Pipilo naufragus [116]

Sp. nov.

Valid

Storrs L. Olson

David B. Wingate

Middle Pleistocene to Holocene

Cave deposits

 Bermuda

A Passerellidae, towhee.

Pliocalcarius orkhonensis [109]

Gen. nov. et Sp. nov

Valid

Nikita V. Zelenkov

Evgeny N. Kurochkin

Late Pliocene

Villafranchian,

Chikoi Formation,

MN 16a

 Mongolia

A probable close relative of longspurs, Calcariidae. The type species of the new genus.

Praealauda hevesensis [95][98][99]

Gen. nov. et Sp. nov.

Valid

Jenö Kessler

János Hír

Middle Miocene

MN 7/8

 Hungary

An Alaudidae. The type species of the new genus.

Primophaps schoddei [117]

Gen. nov. et Sp. nov.

Valid

Trevor H. Worthy

Late Oligocene to early Miocene

Riversleigh

 Australia

 Queensland

A Columbidae related to the bronzewing pigeons. The type species of the new genus.

Pterocnemia mesopotamica [118]

Sp. nov.

Valid

Federico L. Agnolín

Jorge I. Noriega

Late Miocene

Ituzaingó Formation,

Huayquerian

 Argentina

A Rheidae.

Rallicrex litkensis [119]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early Miocene

MN 5

 Hungary

A Rallidae, a new species of Rallicrex Lambrecht, 1933, the type species is Rallicrex kolozsvarensis Lambrecht, 1933.

Rhodospiza shaamarica [109]

Sp. nov.

Valid

Nikita V. Zelenkov

Evgeny N. Kurochkin

Late Pliocene

Villafranchian,

Chikoi Formation,

MN 16a

 Mongolia

A Fringillidae, Carduelinae.

Sanshuiornis zhangi [120]

Gen. nov. et Sp. nov.

Valid

Wang Min

Gerald Mayr

Zhang Jiangyong

Zhou Zhonghe

Middle Eocene

Huayong Formation

 China

A Ciconiiformes Incertae Sedis.

Schizooura lii [121]

Gen. nov. et Sp. nov

Valid

Zhou Shuang

Zhou Zhonghe

Jingmai K. O’Connor

Aptian

Jiufotang Formation

 China

A basal Ornithuromorpha. The type species of the new genus.

Scolopacimilis lartetianus [103]

Gen. nov. et Comb. nov.

Valid

Vanessa L. De Pietri

Mayr

Early Miocene

MN 2a

 France

A Scolopacidae, a new genus for the species "Totanus" lartetianus, the type species of the genus, creating a Comb. nov.

Shengjingornis yangi [122]

Gen. nov et Sp. nov.

Valid

Li Li

Wang Jinqi

Zhang Xi

Hou Shilin

Early Cretaceous

Jiufotang Formation

 China

A Longipterygidae Zhang, Zhou, Hou L. H. et Gu, 2000, Enantiornithes Walker, 1981. The type species of the new genus.

Songzia acutunguis [123]

Sp. nov.

Valid

Wang Min

Gerald Mayr

Zhang Jiangyong

Zhou Zhonghe

Early Eocene

Yangxi Formation

 China

A Songziidae Hou, 1990, Gruiformes. The type species is Songzia heidangkouensis Hou, 1990.

Sturnus kretzoii [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Late Miocene

MN 9

 Hungary

A Sturnidae.

Tadorna minor [119]

Sp. nov.

Valid

Jenö Kessler

János Hír

Miocene

MN 7/8

 Hungary

An Anatidae.

Turdicus minor [98][99]

Sp. nov.

Valid

Jenö Kessler

János Hír

Early to Middle Miocene

 Hungary

A Turdidae.

Tydea septentrionalis [124]

Gen. nov. et Sp. nov.

Valid

Gerald Mayr

Thierry Smith

Early Oligocene

Rupelian

 Belgium:

 Antwerp

An Diomedeidae. The type species of the new genus.

Xiangornis shenmi [125]

Gen. nov. et Sp. nov.

Valid

Hu Dongyu

Xu Xing

Hou Lianhai

Corwin Sullivan

Early Cretaceous

Jiufotang Formation

 China

An Enantiornithes Walker, 1981. The type species of the new genus.

Newly named pterosaurs

Name Novelty Status Authors Age Unit Location Notes Images

Aerotitan[126]

Gen. et sp. nov

Valid

Novas et al.

Late Cretaceous

 Argentina

An azhdarchid. The type species is Aerotitan sudamericanus.

Bellubrunnus[127]

Gen. et sp. nov

Valid

Hone et al.

Late Kimmeridgian

Pfraundorf-Heitzenhofen Basin

 Germany

A rhamphorhynchid. The type species is Bellubrunnus rothgaengeri.

Bellubrunnus.

Dendrorhynchoides mutoudengensis[128]

Sp. nov

Valid

Lü & Hone

Middle Jurassic

Tiaojishan Formation

 China

An anurognathid, a species of Dendrorhynchoides.

Europejara[129]

Gen. et sp. nov

Valid

Vullo et al.

Upper Barremian

La Huérguina Formation

 Spain

A tapejarid. The type species is Europejara olcadesorum.

Europejara.

Gladocephaloideus[130]

Gen. et sp. nov

Valid

Lü, Ji, Wei, & Liu

Barremian

Yixian Formation

 China

A gallodactylid ctenochasmatoid.

Guidraco[131]

Gen. et sp. nov

Valid

Wang, Kellner, Jiang, & Cheng

Early Cretaceous

Jiufotang Formation

 China

A close relative of Ludodactylus. The type species is Guidraco venator.

Guidraco.

Jianchangnathus[132]

Gen. et sp. nov

Valid

Cheng, Wang, Jiang, & Kellner

Middle Jurassic

Tiaojishan Formation

 China

A scaphognathid.

Moganopterus[133]

Gen. et sp. nov

Valid

et al.

Early Cretaceous

Yixian Formation

 China

A boreopterid. The type species is Moganopterus zhuiana.

Qinglongopterus[134]

Gen. et sp.

Valid

Lü, Unwin, Zhao, Gao & Shen.

Middle Jurassic

Tiaojishan Formation

 China

A rhamphorhynchid.

Archosauria incertae sedis

Newly named archosaurs with uncertain affinities

Name Novelty Status Authors Age Unit Location Notes Images

Samrukia[135]

Gen. et sp. nov

Valid

Naish, Dyke, Cau, Escuillié, Godefroit

Late Cretaceous

Kyzylorda District

 Kazakhstan

A possible basal ornithuromorph;[135] alternatively, it could be a large pterosaur.[136][137]

Smok[138]

Gen. et sp. nov

Valid

Niedźwiedzki, Sulej & Dzik

Latest Norian/Early Rhaetian

Lipie Śląskie clay-pit

 Poland

A large predatory archosaur of uncertain phylogenetic placement, the largest known predatory archosaur from the Late Triassic and Early Jurassic of central Europe.

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