Neosuchia

Neosuchia
Temporal range: Early Jurassic - Recent, 196.5–0 Ma
Goniopholis
Scientific classification
Kingdom: Animalia
Phylum: Chordata
Class: Reptilia
Superorder: Crocodylomorpha
Clade: Metasuchia
Clade: Neosuchia
Suborders & Families

See text

Neosuchia is a clade within Mesoeucrocodylia that includes all modern extant crocodilians and their closest fossil relatives.[1] It is defined as the most inclusive clade containing all crocodylomorphs more closely related to Crocodylus niloticus (the Nile Crocodile) than to Notosuchus terrestris.[2] Neosuchia is very diverse and may be polyphyletic, as the clade has undergone many revisions since it was first named in 1988.[3] Neosuchians first appear in the Early Jurassic with the earliest known goniopholid Calsoyasuchus, which lived during the Sinemurian and Pliensbachian stages.[4]

Characteristics

A tooth notch between the maxilla and premaxilla is a basal characteristic of the Neosuchia, although it is lost in some more derived forms, most notably alligatorids.

Classification

Below is a cladogram showing the phylogenetic relationships of neosuchians from Buscalioni et al., 2011:[5]

Neosuchia

Theriosuchus




Goniopholis




Bernissartia fagesii




Susisuchus anatoceps




Las Hoyas neosuchian


Eusuchia

Isisfordia duncani




Hylaeochampsidae

Hylaeochampsa vectiana




Iharkutosuchus




Pietraroiasuchus ormezzanoi



Pachycheilosuchus trinquei







Allodaposuchus sp.



Allodaposuchus precedens




Crocodylia



Borealosuchus sternbergii




Borealosuchus formidabilis




Borealosuchus wilsoni



Borealosuchus acutidentatus





Gavialoidea

Eothoracosaurus mississippiensis




Thoracosaurus macrorhynchus




Thecachampsoides minor




Eogavialis africanum




Gavialis gangeticus




Siquisiquesuchus venezuelensis



Gryposuchus colombianus









Brevirostres

Pristichampsus



Alligatoroidea

Leidyosuchus canadensis





Diplocynodon darwinii



Baryphracta deponiae





Brachychampsa montana




Stangerochampsa mccabei





Alligator mississippiensis




Paleosuchus trigonatus




Caiman yacare




Caiman latirostris



Melanosuchus niger








Navajosuchus mooki



Hassiacosuchus haupti








Crocodyloidea

Asiatosuchus germanicus



Prodiplocynodon langi




Crocodylus affinis




Brachyuranochampsa eversolei



Crocodylus acer





Australosuchus clarkae




Crocodylus megarhinus





Crocodylus cataphractus



Crocodylus porosus





Osteolaemus tetraspis



Crocodylus robustus








Crocodylus spenceri




Gavialosuchus americanus




Tomistoma schlegelii



Tomistoma lusitanica



















In 2012, paleontologists Mario Bronzati, Felipe Chinaglia Montefeltro, and Max C. Langer conducted a broad phylogenetic analysis to produce supertrees of Crocodyliformes, including 184 species. The most parsimonious trees were highly resolved, meaning the phylogenetic relationships found in the analysis were highly likely. Below is a consensus tree from the study:[6]

Neosuchia

Khoratosuchus jintasakuli



Stolokrosuchus lapparenti





Pachycheilosuchus trinquei



Montsecosuchus depereti




Atoposaurus



Alligatorellus beaumonti





Alligatorium




Theriosuchus sympiestodon



Theriosuchus pusillus



Theriosuchus grandinaris






Goniopholididae


Eutretauranosuchus delfsi




Calsoyasuchus valliceps




Sunosuchus miaoi



Sunosuchus junggarensis







Vectisuchus leptognathus




Siamosuchus phuphokensis



Goniopholis stovalli



Goniopholis simus



Goniopholis lucasii



Goniopholis crassidens



Goniopholis baryglyphaeus







Laganosuchus thaumastos




Bernissartia fagesii




Leidyosuchus canadensis



Las Hoyas Neosuchia



Glen Rose Form





Shamosuchus djadochtaensis



Rugosuchus nonganensis






Susisuchus jaguaribensis



Susisuchus anatoceps



Eusuchia

Isisfordia duncani





Allodaposuchus precedens




Iharkutosuchus makadii



Hylaeochampsa vectiana







Crocodylia



Pristichampsus vorax





Borealosuchus wilsoni



Borealosuchus sternbergii



Borealosuchus formidabilis



Borealosuchus acutidentatus
















Elosuchus





Pholidosaurus purbeckensis



Pholidosaurus decipiens






Terminonaris robusta



Sarcosuchus imperator



Sarcosuchus hartti



Dyrosauridae

Oceanosuchus boecensis



Sokotosuchus ianwilsoni




Chenanisuchus lateroculi




Phosphatosaurus gavialoides




Cerrejonisuchus improcerus





Dyrosaurus phosphaticus



Dyrosaurus maghribensis





Arambourgisuchus khouribgaensis




Hyposaurus rogersii




Congosaurus bequaerti




Guarinisuchus munizi




Rhabdognathus sp.



Atlantosuchus coupatezi




Rhabdognathus keiniensis



Rhabdognathus aslerensis
















Thalattosuchia

Pelagosaurus typus




Metriorhynchoidea




"Mystriosaurus"



Platysuchus multiscrobiculatus




Steneosaurus larteti



Steneosaurus edwardsi



Steneosaurus durobrivensis



Steneosaurus boutilieri



Steneosaurus bollensis



Machimosaurus hugii




Steneosaurus priscus



Steneosaurus baroni





Steneosaurus leedsi



Steneosaurus heberti





Steneosaurus gracilirostris



Steneosaurus brevior




Steneosaurus pictaviensis



Peipehsuchus teleorhinus




Teleorhinus calvadosii



Steneosaurus megarhinus











See also

References

  1. Wilson, J. A.; Malkani, M. S.; Gingerich, P. D. (2001). "New crocodyliform (Reptilia, Mesoeucrocodylia) from the Upper Cretaceous Pab Formation of Vitakri, Balochistan (Pakistan)" (PDF). Contributions from the Museum of Paleontology, University of Michigan. 30 (12): 321–336. Retrieved 2009-11-02.
  2. Larsson, H. C. E.; Sidor, C. A.; Gado, B.; Gado, B (2001). "The giant crocodyliform Sarcosuchus from the Cretaceous of Africa" (PDF). Science. 294 (5546): 1516–1519. doi:10.1126/science.1066521. PMID 11679634. Retrieved 2009-11-02.
  3. Benton, M. J. & Clark, J. M. (1998). Archosaur phylogeny and the relationships of the Crocodylia, p. 289-332. In Benton, M.J. (ed.), The Phylogeny and Classification of Tetrapods, Vol. 1 ., Clarendon Press, Oxford.
  4. Tykoski, R. S.; Rowe, T. B.; Ketcham, R. A.; Colbert, M. W. (2002). "Calsoyasuchus valliceps, a new crocodyliform from the Early Jurassic Kayenta Formation of Arizona" (PDF). Journal of Vertebrate Paleontology. 22 (3): 593–611. doi:10.1671/0272-4634(2002)022[0593:CVANCF]2.0.CO;2. Retrieved 2009-11-02.
  5. Buscalioni, A.D.; Piras, P.; Vullo, R.; Signore, M.; Barbera, C. (2011). "Early eusuchia crocodylomorpha from the vertebrate-rich Plattenkalk of Pietraroia (Lower Albian, southern Apennines, Italy)". Zoological Journal of the Linnean Society. 163: S199–S227. doi:10.1111/j.1096-3642.2011.00718.x.
  6. Bronzati, M.; Montefeltro, F. C.; Langer, M. C. (2012). "A species-level supertree of Crocodyliformes". Historical Biology: 1. doi:10.1080/08912963.2012.662680.
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