Temporal range: 318–176 Ma

Middle Carboniferous - Middle Jurassic

Scientific classification
Kingdom: Animalia
Phylum: Arthropoda
Class: Insecta
Superorder: Endopterygota
Order: Miomoptera
Martynov, 1928[1]
  • Metropatoridae
  • Archaemiopteridae
  • Palaeomanteidae


Miomoptera is an extinct order of insects. Although it is thought to be a common ancestor of all holometabolous insects, because no smooth transition between Miomoptera and other holometabolous insect orders is known it is considered to be in a separate order unto itself.

The Miomopterans were small insects, with unspecialised chewing mandibles and short abdominal cerci. They had four wings of equal size, with a relatively simple venation, similar to that of the more primitive living holometabolous insects, such as lacewings.[2]

Adult morphology suggests the adults lived in open habitats. The morphology and gut content shows they fed on the pollen and strobili of gymnosperms. Based on the morphology of the ovipositor, larvae also fed on the pollen of strobili, moving between the scales from one microsporangium to another.[3]

Families and genera



  1. 1 2 Storozhenko, S. Yu.; V. G. Novokshonov (August 1999). "To the Knowledge of the Fossil Family Permosialidae (Insecta: Miomoptera)" (PDF). Far East Entomologist. 76: 1–5. ISSN 1026-051X. Retrieved 2008-06-12.
  2. Hoell, H.V., Doyen, J.T. & Purcell, A.H. (1998). Introduction to Insect Biology and Diversity, 2nd ed. Oxford University Press. p. 322. ISBN 0-19-510033-6.
  3. Novokshonov, V. G.; L. V. Zhuzhgova (2004). "Discussion of the System and Phylogeny of the Order Palaeomanteida (= Miomoptera) (...)" (PDF). Paleontological Journal. 38 (Suppl. 2): S173–S184. Retrieved 2008-04-11.
  4. Tree of Life Web Project. 2003. Miomoptera. in The Tree of Life Web Project,
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