Temporal range: Miocene–Recent
|A live individual of the Spanish slug Arion vulgaris, in the wild|
|(unranked):|| clade Heterobranchia|
Tetraspididae Hagenmüller, 1885
Unlike some slugs, European Arionidae have no internal shell. They also have no keel on the back. The caudal mucous pit is above the tip of the tail. The respiratory pore (pneumostome) is in front of the midpoint of the mantle. The body length is up to 250 mm. Mantle covers only a part of the body and lies in the anterior part.
The jaw is odontognathic, which means it is transversally ribbed. Radular teeth include: central tricuspid, lateral bi- or tricuspid, marginal bicuspid, all having broad bases. Teeth are often accreted. The digestive system forms 2 loops. The heart, in relation to body axis, is titled to the left. The kidney is circular (surrounding aorta). Cephalic retractors tend to divide into separate branches attached independently to the posterior part of pallial complex. The shell is strongly reduced in places, most often completely buried in the mantle, usually of loose crystals or plate-like. Genitalia: the penis is present only in some species, epiphallus is present in nearly all of them. Male copulatory organs are generally reduced, their role being taken over by a well-developed atrium and the epiphallus that produces spermatophores.
Family Arionidae has no subfamilies according to the taxonomy of the Gastropoda by Bouchet & Rocroi, 2005.
Genera within the family Arionidae include:
- Anadenulus Cockerell, 1890
- Ariolimax Mörch, 1859 - Banana slug
- Arion Férussac, 1819
- Ariunculus Lessona, 1881
- Carinacauda Leonard, Chichester, Richart & Young, 2011
- Geomalacus Allman, 1843
- Gliabates Webb, 1959
- Hemphillia Bland & Binney, 1872
- Hesperarion Simroth, 1891
- Letourneuxia Bourguignat, 1866
- Magnipelta Pilsbry, 1953
- Prophysaon Bland & W. G. Binney, 1873
- Securicauda Leonard, Chichester, Richart & Young, 2011
- Udosarx Webb, 1959
- Zacoleus Pilsbry, 1903
The parasites of the Arionidae slugs include the Sciomyzidae.
- "Family summary for Arionidae". AnimalBase, last change 12-06-2009, accessed 4 August 2010.
- Barker G. M.: Gastropods on Land: Phylogeny, Diversity and Adaptive Morphology. in Barker G. M. (ed.): The biology of terrestrial molluscs. CABI Publishing, Oxon, UK, 2001, ISBN 0-85199-318-4. 1-146, cited pages: 139 and 142. (In the reference specified as Ariononae.)
- Cockerell (1890). Ann. Mag. nat. Hist. (6)6: 278.
- Leonard, William P.; Chichester, Lyle; Richart, Casey H.; Young, Tiffany A. (2011). "Securicauda hermani and Carinacauda stormi, two new genera and species of slug from the Pacific Northwest of the United States (Gastropoda: Stylommatophora: Arionidae), with notes on Gliabates oregonius Webb 1959" (PDF). Zootaxa. 2746: 43–56.
- Webb (1959). Gastropodia 1(3): 22.
- Bland T. & Binney W. G. (1874). "Description of Hemphillia, a new Genus of Terrestrial Mollusks". Annals of the Lyceum of Natural History of New York. 10: 208-211. Plate 9.
- Simroth (1891). Malak. Bl. (N.S.) 11: 111.
- Pilsbry H. A. (1953). Nautilus 67: 37.
- Bland & Binney W. G. (1873). Ann. Lyceum nat. Hist. N. York 10(3): 921.
- Webb (1959). Gastropodia 1: 22.
- Pilsbry H. A. (1903). Proc. Acad. nat. Sci. Philad. 55: 626.
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